| PI | Institution | Founded / Current Sites |
| Peter Adler | Utah State University | Sheep Experimental Station |
| Elizabeth Borer | University of Minnesota | Bunchgrass (Andrews LTER), Buttercup (Andrews LTER), Cedar Creek LTER, Finley NWR, Hastings UCNRS, Hopland REC, Lookout (Andrews LTER), Mclaughlin UCNRS, Sierra Foothills REC, Sedgwick Reserve UCNRS |
| Jennifer Firn | Queensland University of Technology | Burrawan |
| Daniel Gruner | University of Maryland | Sagehen Creek UCNRS |
| W. Stanley Harpole | Iowa State University | Chichaqua Bottoms, Cedar Creek LTER, Hopland REC, Mclaughlin UCNRS, Sedgwick Reserve UCNRS, Sierra Foothills REC |
| Eric Lind | University of Minnesota | NutNet Databases |
| John Orrock | University of Wisconsin-Madison | Savannah River, Tyson |
| Eric Seabloom | University of Minnesota | Bunchgrass (Andrews LTER), Buttercup (Andrews LTER), Cedar Creek LTER, Finley NWR, Hastings UCNRS, Hopland REC, Lookout (Andrews LTER), Mclaughlin UCNRS, Sierra Foothills REC, Sedgwick Reserve UCNRS |
| Melinda Smith | Yale University | Konza LTER, Saline Experimental Range |